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Shiro TSUYUZAKI
Plant community ecology / Environmental conservation

Mount Usu / Sarobetsu post-mined peatland
From left: Crater basin in 1986 and 2006. Cottongrass / Daylily

(Update on Feb. 20 2010)

Rough cocklebur
Xanthium occidentale Bertoloni [Asteraceae]

O˙onamomi (オオオナモミ, 大葈耳)

Lifeform: annual forb
Distribution: origin in the north America (exotic to Japan)
Habitat: open fields in urbanized regions
Seed dispersal: external animal
Poisonous plant


Xanthium occidentale in Hokkaido

X. occidentale is native and the others are exotic.

X. italicum Moretti (イガオナモミ)
X. occidentale Bertoloni (オオオナモミ)
X. spinosum L. (トゲオナモミ)
X. strumarium L. (オナモミ)


Xanthium occidentale in Hokkaido

ST ST ST
[Left] A shoot. [Center] close-up of fruits. [Rgiht] close-up of leaf. At N25/W3, North Ward, Sapporo, on August 30 2010.


References

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