(First upload on May 5 2009. Last on March 27 2017) [ 日本語 | English ]

Sambucus racemosa L.

Mount Usu / Sarobetsu post-mined peatland
From left: Crater basin in 1986 and 2006. Cottongrass / Daylily

ssp. racemosa (セイヨウアカミニワトコ), the type in North America and Europe
ssp. sieboldiana (Miq.) Hara (ニワトコ, 接骨木/庭常)
Distribution: China - southern Korea - Japan (not in Hokkaido)
Habitat: common on mountain hills
Stem: 3-6 m tall
Dormant bud: sharp top

var. major (Nakai) Murata (オオニワトコ)
f. aurantiaca (Nakai) Hara (ダイダイミノニワトコ)
f. dissecta Murata (サケバニワトコ)
f. glaberrima (Nakai) Hara ex Honda (ケナシニワトコ)
f. macropylla Hara (オオバニワトコ)

f. nakaiana Murata (キミノニワトコ)
f. ovatifolia Hara (マルバニワトコ)
f. stenophylla (Nakai) Hara (ケオオニワトコ)

ssp. kamtschatica (E. L. Wolf) Hultén
Ezoniwatoko (エゾニワトコ, 蝦夷接骨木), red elder
Lifeform: deciduous shrub
Distribution: Russian Far East - Korea - Japan (Hokkaido - northern Honshu)
Habitat: woodlands - forests

HUSTEP: Plants and Plant communities in Japan (flora list)

Seed dispersal: animal
Leaf: dense > ssp. sieboldiana
Inflorescence: granular process → extended, hair-like
Chromosome number: 2n = 38

Sambucus sieboldiana (Miq.) Blume ex Graebn. var. miquelii (Nakai) H. Hara, excl. basionym
Sambucus racemosa L. var. miquelii auct. non Nakai
Sambucus kamtschatica E. L. Wolf
f. aureocarpa (Hara) Hara ex Honda (キミノエゾニワトコ)
f. hiraii (Miyabe et Tatewaki) Hara ex Honda (ホソバエゾニワトコ)
f. nakayamai (Yanagita) Hara ex Honda (ビロードエゾニワトコ)
f. nuda Hara (ケナシエゾニワトコ)
f. platyphylla (Miyabe et Tatewaki) Hara ex Honda (ヒロハエゾニワトコ)

⇒ APG: Adoxaceae

Sambucus L. (ニワトコ) in Japan

Lifeform: short tree - shrub
Distribution: one species (with two varieties) in Hokkaido
S. canadensis L. (アメリカニワトコ)
S. chinensis Lindl. (ソクズ)

var. formosana (Nakai) H. Hara (タイワンソクズ)

S. nigra L. (セイヨウニワトコ)
S. racemosa L. (ニワトコ)
S. williamsii Hance (コウライニワトコ)

Sambucus racemosa L. in HU campus

ssp. sieboldiana

[1-3] in the Koishikawa Botanical Garden (小石川植物園), the University of Tokyo, on March 16 2017.

ssp. kamtschatica



[1/2] at the backyard of the university co-op central restaurant on April 29 2008. [3/4] at the backyard of the Institute of Low Temperature Science, Hokkaido University, Sapporo, on May 19 2016. [5] fruits in Genshirin on July 23 2009. Discarded: inflorescences found in Genshirin (原始林) or Keiteki-no-mori Forest (恵迪の森) on May 19 2008.