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(First upload on January 8 2012. Last on November 13 2018) [ 日本語 | English ]

Bidens L. (センダングサ) Kosendangusa (コセンダングサ, 小栴檀草), cobbler's pegs or Spanish needle Lifeform: weedy annual forb Distribution: extotic worldwide, including Japan (probably originating from South Africa) Habitat: ruderal, such as roadsides

Seed dispersal: animal (external) by hooked barbs Involucral segment: short ↔ B. frondosa: long Chromosome number: 2n = 24, 36, 48, or 72 var. bisetosa S. Ohtani et S. Suzuki (アワユキセンダングサ) var. minor (Blume) Sherff (コシロノセンダングサ/シロノセンダングサ), developing white ligulate flowers var. pilosa (コセンダングサ, s.s.), without ligulate flowers var. radiata Sch. Bip. (タチアワユキセンダングサ)

Bidens pilosa in Japan

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var. pilosa

1 2 3
4 5 6
var. pilosa. [1-3] at Yodogawa Riverside Park, Osaka, western Japan, on October 29 2011. [4/5] along Sophia Street, Chiyoda Ward, Tokyo, on November 14 2014. [6] near a cemetery (Koumyoujoen (光明浄苑), Machida City, Tokyo, on December 10 2017.

var. minor

1 2 3
[1-3] on a bank along Sophia Street, Chiyoda Ward, Tokyo, on October 19 2018.