• HOME
  1. 日本語
  2. English
• Personal Info.
  1. CV
  2. For candidates
  3. Give me job!
  4. Recent(?) Photos
  5. Master/doctor theses
• Education
  1. Catalog
  2. HUSTEP
  3. Ecology
  4. Natural env. studies
  5. Environ. conserv.
  6. Field trip for Env. Res.
     
  7. Eco-campus(plants)
  8. Symbiosis in nature
  9. Beadroll
  10. Glossary(References)
• Research
  1. Abstract
  2. Volcanic succession
  3. Wildfire ecology
     
  4. Wetland conservation
  5. Skislope vegetation
  6. Mining
  7. Pub: English
  8. Pub: Japanese
• Hobby · Link
  1. Yoshida
  2. Plant taxonomy
  3. Plants
  4. Cup of tea
  5. Western Australia
  6. Link
  7. Contact

(First upload on February 12 2011. Last on January 3 2026) [ 日本語 | English ]

[Engler's syllabus エングラー体系]

Humulus L. (カラハナソウ) in Japan

---

H. scandens (Lour.) Merr. (カナムグラ)

H. lupulus L. (カラハナソウ)

Humulus lupulus in Hokkaido

---

Distribution: temperate Northern Hemisphere, including northern Japan) var. cordifolius (Miq.) Maxim. ex Franch. et Sav. Karahanasou (カラハナソウ, 唐花草), Japanese hop Life form: dioecious, perennial climber Distribution: Eastern Asia Habitat: grassland - forest Field training on integrated environmental research (統合環境調査法実習) (fauna and flora 種リスト) Hair: much on leaves and stems

Spine: short, stiff spines, giving a rough texture var. lupuloides E. Small (syn. H. americanus Nutt.): Eastern North America var. lupulus: Europe - western Asia, exotic to Japan (セイヨウカラハナソウ/ホップ), common hop Hair: less on leaves and stems Spine: few or inconspicuous on petioles var. neomexicanus A. Nelson et Cockerell: Western North America var. pubescens E. Small: Midwest North America

var. cordifolius (カラハナソウ)

1 2 3
[1-3] in Toya Lake Experimental Station, Hokkaido University, on August 24 2010. It was rain.

4 5 6
[4-6] on the banks of the Honkawa River, a tributary of the Satsunai River, on June 23 2025.

7 8 9
[7/8] along a forest road in Shimokawa Town, northern Hokkaido, on May 27 2016. [9] in a forest at the backyard of Toya Lake Experimental Station on September 13 2016. Records:

Mount Usu

1 2 3
[1-3] near the Nishiyama Crater Group at the foot of Mount Usu on September 18 2013.