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(First upload on September 27 20120. Last on December 29 2021) [ 日本語 | English ]

Galium L. (ヤエムグラ) Togenashimugura (トゲナシムグラ, 刺無葎), wild madder, white bedstraw, upright bedstraw, whorled bedstraw or hedge bedstraw Lifeform: perennial forb, developing grassy thicket, with 15-100 cm in height less erect with ascending branches Distribution: naturally in Europe (exotic to Japan and North America) Habitat: road sides, river banks, etc.

Leaf: whorls of six to eight leaves Stem: with no spines and with four ridges Chromosome number: 2n = 22 Synonyms: Galium erectum Huds. Galium mollugo ssp. erectum (Huds.) Briq. Galium mollugo var. erectum (Huds.) Domin Rubia mollugo (L.) Baillon

Galium mollugo in Hokkaido

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[1-4] along the formerly Shiretoko Forest Road in eastern Hokkaido on August 29 2020. [5] around the accomodation of Teshio Experimental Forest, northern Hokkaido, on August 30 2021. [6] along a paved road in Utasai wetland (歌才湿原), southern Hokkaido, on October 16 2024.