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(First upload on September 27 20120. Last on December 29 2021) [ 日本語 | English ]

Galium L. (ヤエムグラ) Togenashimugura (トゲナシムグラ, 刺無葎), wild madder, white bedstraw or upright bedstraw Lifeform: perennial forb, developing grassy thicket Distribution: naturally in Europe (exotic to Japan)

Habitat: road sides, river banks, etc. Stem: no spines with four ridges

Galium mollugo in Hokkaido

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4 5
[1-4] along the formerly Shiretoko Forest Road in eastern Hokkaido on August 29 2020. [5] around the accomodation of Teshio Experimental Forest, northern Hokkaido, on August 30 2021.