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(First upload on July 20 2014. Last on June 10 2019) [ 日本語 | English ]

Paeoniaceae Raf. (ボタン) 33 species in the one genus Paeonia L. (ボタン) Shrub P. suffruticosa Andrews (ボタン) Herb P. japonica (Makino) Miyabe et Takeda (ヤマシャクヤク) P. lactiflora Pall. (シャクヤク, s.l.)

P. obovata Maxim. (ベニバナヤマシャクヤク) f. albiflora M. Mizush. ex T.Shimizu (ケナシシロシャクヤク) f. glabra (Makino) Kitam. (ケナシベニバナヤマシャクヤク) f. oreogeton (S.Moore) Kitag. (マンシュウヤマシャクヤク) P. officinalis L. (オランダシャクヤク) P. tenuifolia L. (ホソバシャクヤク) ⇒ Engler's syllabus (エングラー体系) merged into Ranunclaceae (Ohwi 1983)

Chinese tree peony = eustele = remarkable cambium → medullary ray is present Chinese tree peony → Chinese peony (formation of stamens and the development process Homology of proteins: genealogical relationship is far between Paeoniaceae and Ranunculaceae Anatomy: close to Hypericaceae, Dilleniaceae and Theaceae

Paeonia suffruticosa and P. lactiflora transplanted in Hokkaido

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Paeonia suffruticosa

Botan (ボタン, 牡丹), Chinese tree peony Lifeform: shrub

Distribution: native to northeastern China (extotic to Japan) Leaflet: lacinia

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[1/2] at a house garden in East Ward, Sapporo, on June 2 2014. [3/4] at a house garden in East Ward, Sapporo, on May 20 2019.

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Paeonia lactiflora

Syakuyaku (シャクヤク, 芍薬, s.l.), Chinese peony, or common garden peony
Lifeform: perennial forb
Distribution: native to central to eastern Asia
Leaflet: rounded
var. lactiflora (シベリアシャクヤク)
var. trichocarpa (Bunge) Stearn (シャクヤク, s.s.)

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[1-3] undar a street trees in North Ward, Sapporo, on June 21 2014.