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(Upload on February 11 2023) [ 日本語 | English ]

Rhamnaceae Juss. (クロウメモドキ科)






Mount Usu / Sarobetsu post-mined peatland
From left: Crater basin in 1986 and 2006. Cottongrass / Daylily

[Engler's syllabus (エングラー体系)]
Order Rhamnales (クロウメモドキ)
Family Rhamnaceae Juss. (クロウメモドキ), buckthorn family
Lifeform: trees, shurbus and vines
Distribution: worldwide, more in subtropical and tropical regions
APG: Order Rosales (バラ)
950 species in 55 genera
Tribe Ampelozizipheae
Ampelozizyphus Ducke
Tribe Bathiorhamneae
Bathiorhamnus Capuron
Tribe Colletieae
Adolphia Meisn.
Colletia Comm. ex Juss.
Discaria Hook.
Kentrothamnus Suess. et Overkott
Ochetophila Poepp. ex Reissek
Retanilla (DC.) Brongn.
Trevoa Miers ex Hook.
Tribe Doerpfeldieae
Doerpfeldia Urb.
Tribe Gouanieae
Alvimiantha Grey-Wilson
Crumenaria Mart.
Gouania Jacq.
Helinus E. Mey. ex Endl.
Johnstonalia Tortosa (originally named Johnstonia; by some treated as Gouania)
Pleuranthodes Weberb. – synonym of Gouania
Reissekia Endl.
Tribe Maesopsideae
Maesopsis Engl.
Tribe Paliureae
Condaliopsis (Weberb.) Suess.
Hovenia Thunb.
Paliurus Mill.
Ziziphus Mill. (ナツメ)
Sarcomphalus P. Browne emend. Hauenschild
Tribe Phyliceae
Nesiota Hook.f., extinct
Noltea Rchb.
Phylica L.
Trichocephalus Brogn.
Tribe Pomaderreae
Blackallia C. A. Gardner
 Cryptandra Sm.
Papistylus Kellermann, Rye et K. R. Thiele
Polianthion K. R. Thiele
Pomaderris Labill.
Serichonus K. R. Thiele
Siegfriedia C. A. Gardner
Spyridium Fenzl
Stenanthemum Reissek
Trymalium Fenzl
Tribe Rhamneae
Auerodendron Urb.
Berchemia Neck. ex DC.
Berchemiella Nakai (ヨコグラノキ)
B. berchemiifolia (Makino) Nakai (ヨコグラノキ)
Condalia Cav.
Frangula Mill.
Karwinskia Zucc.
Krugiodendron Urb.
Phyllogeiton (Weberb.) Herzog
Pseudoziziphus Hauenschild
Reynosia Griseb.
Rhamnella Miq.
Rhamnidium Reissek
Rhamnus L. (クロウメモドキ)
Sageretia Brongn.
Scutia (Comm. ex DC.) Brongn.
Tribe Ventilagineae
Smythea Seem.
Ventilago Gaertn.

ST
Ceanothus velutinus

Incertae sedis
Alphitonia Endl., close to Colubrina
Araracuara Fern. Alonso
Ceanothus L., close to Pomaderreae
Chaydaia Pit., syn. Rhamnella
Colubrina Rich. ex Brongn.
Conalma G. L. Nesom
Emmenosperma F. Muell., close to Colubrina
Fenghwaia G. T. Wang et R. J. Wang
Granitites Rye, close to Alphitonia
Hybosperma Urb., syn. Colubrina
Jaffrea H. C. Hopkins et Pillon
Lasiodiscus Hook.f.
Oreorhamnus Ridl.
Schistocarpaea F. Muell., prob. belonging to Colletieae

Rhamnus L. (クロウメモドキ)

R. arguta Maxim. (ウメバクロツバラ)
R. chugokuensis Hatus. (タイシャククロウメモドキ)
R. costata Maxim. (クロカンバ)

f. nambuana (Honda) H. Hara (ナンブクロカンバ)
f. pubescens Hiyama (ヤマナシクロカンバ)

R. davurica Pall. (クロツバラ, s.l.), Dahurian buckthorn

var. davurica (チョウセンクロツバラ)
var. nipponica Makino (クロツバラ)

f. pubescens H. Hara (ケクロツバラ)

R. diamantiaca Nakai (イヌクロウメモドキ)
R. formosana Matsum. (シマクロウメモドキ)
R. ishidae Miyabe et Kudô (ミヤマハンモドキ)
R. japonica Maxim. (クロウメモドキ, s.l.), Japanese buckthorn

var. decipiens Maxim. (クロウメモドキ)

f. chlorocarpa (Honda) H. Hara (アオミノクロウメモドキ)
f. senanensis (Koidz.) H. Hara (シナノクロウメモドキ)

var. japonica (エゾノクロウメモドキ)
var. microphylla H. Hara (コバノクロウメモドキ)

R. kanagusukui Makino (ヒメクロウメモドキ)
R. koraiensis C. K. Schneid. (マルバクロウメモドキ)
R. liukiuensis (E. H. Wilson) Koidz. (リュウキュウクロウメモドキ)
 R. nakaharae (Hayata) Hayata (ナカハラクロウメモドキ)
R. oiwakensis Hayata (コゴメバクロウメモドキ)
R. parvifolia Bunge (イワクロウメモドキ)
R. shozyoensis Nakai (ナガミノクロツバラ)
R. taquetii (H. Lév.) H. Lév. (サイシュウクロツバラ)
R. utilis Decne. (シーボルトノキ), Chinese buckthorn
R. yoshinoi Makino (キビノクロウメモドキ)
R. × calcicola Hatus. (クニガミクロウメモドキ)

Frangula Mill. (イソノキ)

when separated
F. crenata (Siebold et Zucc.) Miq. (イソノキ), syn. R. crenata Siebold et Zucc.

Ziziphus Mill. (ナツメ)

Lifeform: consisting of ca. 40 species of spiny shrubs and small trees
 Distribution: warm-temperate, subtropical and tropical regions
 Z. jujuba Mill. (ナツメ, s.l.)
Z. mauritiana Lam. (イヌナツメ)
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