• HOME
  1. 日本語
  2. English
• Personal Info.
  1. CV
  2. For candidates
  3. Give me job!
  4. Recent(?) Photos
  5. Master/doctor theses
• Education
  1. Catalog
  2. HUSTEP
  3. Ecology
  4. Natural env. studies
  5. Environ. conserv.
  6. Field trip for Env. Res.
     
  7. Eco-campus(plants)
  8. Symbiosis in nature
  9. Beadroll
  10. Glossary(References)
• Research
  1. Abstract
  2. Volcanic succession
  3. Wildfire ecology
     
  4. Wetland conservation
  5. Skislope vegetation
  6. Mining
  7. Pub: English
  8. Pub: Japanese
• Hobby · Link
  1. Yoshida
  2. Plant taxonomy
  3. Plants
  4. Cup of tea
  5. Western Australia
  6. Link
  7. Contact

(First upload on November 21 2011. Last on July 30 2018) [ 日本語 | English ]

⇒ Engler's syllabus (エングラー体系)

Trapaceae L. (ヒシ)

---

Flower: hermaphrodite, radial symmetry, tetramerous choripetalae Evolution Trapella (ヒシモドキ) ↑____Hemitrapa (アスナロビシ) ↑____extinct in Pliocene ↑____discovered two deformed carpels These two genra hve not been discovered in the New Continent ↑ affinity of aquatic life Saxifragaceae Chrysosplenium: warm-temperate Asia

Trapa: most species in Asian Continent → Asian origin One genus (the taxonomy is controversial) 30 species in the world (2-3 species in Japan) Annual aquatic forb (+ two genera in Nymphaeaceae 1. floating leaf = 3-6 cm in diameter, dense hair on the veins of backside 2. two spines on the drupe ___ T. japonica 2. four spines on the drupe ___ T. natans L. var. rubeola Makino (メビシ, not in Hokkaido)

1. floating leaf = 1-2 cm in diameter, nearly no hair on the backside ___ T. incisa Sieb. et Zucc. Trapa L. (ヒシ) T. bicornis Osbeck (コウモリビシ) var. bispinosa (Roxb.) Nakano (トウビシ) var. taiwanensis (Nakai) Z. T. Xiong (タイワンビシ) T. incisa Siebold et Zucc. (ヒメビシ) T. japonica Flerow (ヒシ) T. macropoda Miki (シリブトビシ) T. natans L. (オニビシ) T. pseudincisa Nakai (マンセンビシ)

Trapa japonica in Hokkaido

---

Hishi (ヒシ, 菱), Japanese water chestnut Lifeform: annual aquatic forb Distribution: China - Taiwan - Korea - Japan Habitat: waterlogged wetlands, such as ponds and swamps → wetland (湿原) Field training on integrated environmental research (統合環境調査法実習) (fauna and flora 種リスト) Chromosome number: 2n = 96 Edible plant Synonyms Trapa japonica Flerow var. tuberculifera (V. Vassil.) Tzvelev

Trapa natans L. var. bispinosa Makino, excl. basion. Trapa japonica Flerow var. makinoi (Nakano) Ohba, comb. nud. Trapa bicornis Osbeck var. iinumae (Nakano) Nakano Trapa bicornis Osbeck var. makinoi (Nakano) Nakano Trapa bicornis Osbeck var. iwasakii (Nakano) Nakano Trapa bispinosa Roxb. var. makinoi Nakano Trapa bispinosa Roxb. var. iinumae Nakano

1 2 3
[1] at Zenibako Coast (銭函海岸), Otaru City, on August 31 2011. [2] in the artificial pond (nicknamed Ono Pond, 大野池), south to the Faculty of Engineer in the campus of Hokkaido University on July 19 2018. [3] in Akanuma Pond, Kushiro Moor, eastern Hokkaido, on June 28 2015.